Clarence A. (Bud) Ryan, one of WSU’s preeminent scientists, died suddenly of a brain aneurysm in October. Ryan pioneered the study of the innate immune response of plants. Prior to his work, plants were assumed to contain protease inhibitors all the time, as a deterrent to being eaten. Ryan discovered instead that plants make the inhibitors in response to an attack. He further showed that an attack on one part of a plant sets off chemical signals that spur production of inhibitors throughout the entire plant. Besides his scientific renown, Ryan was well known around campus for his graciousness—-and his ability on the basketball court.

Clarence "Bud" Ryan on right.
Clarence “Bud” Ryan on right.